Primate hybridization

p-ter points me to a new paper which documents interspecies hybridization in monkeys whose lineages putatively diverged about 3 million years ago. Note that the hybridization follows Haldane's rule: the heterogametic sex (in mammals the males) exhibits sterility while the other sex does not. Whatever genetic incompatibilities built over the period during which the two populations became distinct the less robust sex (males have only one copy of the X chromosome, ergo, sex-linked diseases) naturally exhibits greater breakdown in hybrids. In any case, the story is obviously relevant to Neandertal introgression, since the divergence between moderns and Neandertals is on the order of 1 million to 100,000 years.

Related: Mammalian hybridization potentialities.

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Hybrids are sort-of a mystery in evolutionary biology. In the strictest sense, they shouldn't exist because the offspring are often sterile or reproductively impaired.
We usually think of speciation as a bifurcating process -- a single lineage splitting into two. The relationships of those species can often be determined using DNA sequences. But we know that there are exceptions, like horizontal gene transfer in bacteria.
Wild specimen of the butterfly species, Heliconius heurippa.
For some reason, I have been collecting links to articles involving hybridization. That, on its own, would call for a massive link dump, but a recent news item makes for a nice contrast. First, the hybrids:

Gelada (Theropithecus) and Papio have also hybridized. Different genera, although they are both called baboons. In addition, Papio species arguably constitute a single superspecies.