Prenolepis queen

Is Prenolepis more closely related to Lasius and/or Brachymyrmex than to other formicines, or are the similarities just secondary? I mean, they both have tiny males and tiny workers, and their queens also do look similar.

More closely related to Nylanderia and Paratrechina. Trouble is, most non-parasitc formicine queens look a lot alike, one of the reasons for mainly worker-based ant taxonomy.